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Doctoral Thesis

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Nanotemplates for the combined structural and functional analysis of membrane-associated proteins (2019)
Krupp, Nikolai
Plasma membranes are essential for life because they give cells an identity. Plasma membranes are almost impermeable to fluids and substances. Still, transport between inside and outside needs to be possible. An important transport way is endocytosis. This mechanism relies on membrane-associated proteins that sense and induce curvature to the plasma membrane. However, the physics and structural dynamics behind proteins acting on membranes is not well understood. There is a standard method in vitro to investigate membrane-associated proteins sensing spherical geometries: They are incubated on unilamellar vesicles. This procedure allows to analyze these proteins in their bound state. This approach is inappropriate for GRAF1 (GTPase Regulator Associated with Focal Adhesion Kinase-1), a key player in endocytosis because it senses tubular geometries instead. However, GRAF1 extrudes lipid tubes from vesicles that can be analyzed. Still, this is a limited method because these tubes suffer from inhomogeneity and they do not enable the observation of intermediate and lower concentration binding states. To overcome this issue they can be incubated on pre-tubular structures called nanotemplates. There have been studies using carbon nanotubes and Galactosylceramide lipid tubes as nanotemplates. These approaches require complex chemical modifications or expensive components and they are not necessarily flexible. In this work we present a simple and easy new approach to prepare nanotemplates using Folch lipid mixture. We show on the basis of BPG, a truncate of GRAF1, that our nanotemplates are suitable for Cryo-EM and that it is possible to use IHRSR (Iterative Helical Real Space Reconstruction) to analyze the structure of BPG in its bound state. Moreover, the qualification for Cryo-EM allows to use plunge freezing to interrupt the incubation on our nanotemplates abruptly. This enables the analysis of intermediate binding states to understand the binding process.
Balsaminaceae in Madagascar and Comoro Islands: Systematics, Evolution, and Biogeography (2019)
Rahelivololona, Marie Elisette
In this thesis, I present the results of my studies on taxonomy, systematics, and biogeography of Impatiens (Balsaminaceae) in Madagascar and the Comoro islands. In Chapter 1 I reviewed the literature on taxonomy and classification of Balsaminaceae, on habitat, world distribution, morphology, molecular phylogenetics and infrageneric classification of the genus Impatiens. In Chapters 2-15 (Fischer & Rahelivololona 2002, 2003, 2004, 2007, 2015a, 2015b, 2015c, 2016, Fischer et al. 2004. 2017, 2018a, b submitted, Rahelivololona et al. 2003) I presented the first results of a revision of Balsaminaceae of Madagascar and the Comoro islands including the description of 78 new species. In Chapter 16 (Yuan et al. 2004) we worked on the phylogeny and biogeography of Balsaminaceae inferred from ITS sequences using combined results from molecular phylogenetic and morphological analyses. In Chapter 17 (Rahelivololona et al. 2018) we conducted a phylogeny and assessment of the infrageneric classification of species in the Malagasy Impatiens (Balsaminaceae) with a particular emphasis on taxa collected from Marojejy. Below I summarise the most important findings of each chapter and provide an outlook for future studies. How many species of Impatiens occur in Madagascar and the Comoro islands? To provide additional information on the taxonomic revision of Impatiens in Madagascar and the Comoro islands, the identification of already described species as well as the description of new species was conducted. Based on herbarium specimens from BR, G, K, NEU, P, TAN and on living plants collected during several field trips, 78 new species and 6 nomina nova have been published and another 70 new taxa are already identified. Actually more than 260 species occur in Madagascar and the Comoro islands and all of them are endemic. For each species, a description of the morphology, phenology, ecology and known distribution range was provided. Apart from new taxa, the delimitation of already described species like Impatiens firmula Baker and Impatiens hildebrandtii Baill. could be clarified by studying the types and by observing the variability in the field. Are the groups of Impatiens in Madagascar monophyletic, and what is the systematic position of Trimorphopetalum? Yuan & al. (2004) conducted a molecular phylogenetic study to examine the morphological and karyological evolution, and the historical biogeography of the Balsaminaceae family by using nucleotide sequence data of internal transcribed spacer regions of nuclear ribosomal DNA. The results support the monophyly of the Malagasy endemic section Trimorphopetalum and show that the cleistogamous Impatiens inaperta should be included in the sect. Trimorphopetalum which is the most derived within Impatiens. Therefore, the section Preimpatiens proposed by Perrier de la Bâthie (1934) is paraphyletic. Rahelivololona & al. (2018) provided a phylogenetic study focused on three subdivisions (based on macromorphological characters) proposed by Perrier de la Bâthie (1934). The analysis was done using two nuclear AP3/DEF homologues (ImpDEF1 and ImpDEF2) and the plastid atpB-rbcL spacer to reassess or assess the monophyly of the Malagasy Impatiens, of the sections Preimpatiens (Humblotianae and Vulgare groups) and Trimorphopetalum. A focus was on the species of Impatiens from the Marojejy National Park and of the morphologically variable species I. elatostemmoides, I. “hammarbyoides”, I. inaperta and I. manaharensis, using monophyly as the primary criterion. As results the Malagasy Impatiens are paraphyletic and the section Preimpatiens sensu Perrier de la Bâthie (1934) (= subgen. Impatiens sensu Fischer & Rahelivololona 2002) was not resolved as a monophyletic group. The section Trimorphopetalum sensu Perrier de la Bâthie (1934) (= subgen. Trimorphopetalum sensu Fischer & Rahelivololona 2002), however, was strongly confirmed as a monophyletic lineage (BS: 92; BPP: 1). Neither the Humblotianae group nor the Vulgare group was supported as monophyletic. None of the morphologically variable species appeared to be monophyletic and the sampled species of Impatiens from the Marojejy National Park do also not form a monophyletic group. What are the biogeographical position and the distribution patterns of Impatiens in Madagascar and the Comoro islands? Investigation of the geographical affinities and species distribution of section Impatiens (including Humblotianae group and Vulgare group) and section Trimorphopetalum were conducted and the origin and evolution as well as species richness and endemism were discussed. The isolation, the climate and the complex topography of Madagascar have generated the microhabitats and ecological niches favourable to the diversification of Impatiens species. Impatiens of Madagascar with 260 endemic species is actually the largest genus in Madagascar. Therefore, Madagascar and the Comoro islands are among the most species-rich regions in the world for Impatiens. Future studies In Impatiens on Madagascar, there remain numerous unresolved questions that need to be adressed: • A further study based on a much larger molecular data set and sampling from the entire geographic ranges of Impatiens in Madagascar is needed to retest the monophyly of the different subgenera and sections, as well as a molecular dating of the Malagasy Impatiens. • The study of pollinators as a key for understanding the radiation and species richness is required: Within Impatiens the different shapes of spur are related to pollinators (bees, birds, butterflies and moths). Therefore pollinator observation of specific species need to be done to understand the radiation of species by adaptation and coevolution with these pollinators. A pollination study with a large number of species within section Trimorphopetalum will help to understand the mechanism of complete disappearance of the spur, the shift of pollinators and the evolution of species richness. • The destruction of the natural habitats of Impatiens and the subsequent reduction of humidity in logged area constitute a severe threat for the survival of many species. The conservation and reforestation of vulnerable areas such as Ankaratra, Daraina, Mandraka and Col des Tapia near Antsirabe is required. • In terms of conservation and to mitigate the threat on the genus, a study on the ex-situ-conservation of Malagasy Impatiens species is very important as long as some species are suitable for horticultural purposes (e.g. Impatiens mayae-valeriae, Impatiens emiliae and species with broad red spur). • Finally, the publication of the revision of Impatiens of Madagascar and the Comoro islands will help other botanists to identify the species and will thus increase our knowledge on the group.
Mechanisms of methane bubble formation, storage and release in freshwater sediments (2019)
Liu, Liu
Lakes and reservoirs are important sources of methane, a potent greenhouse gas. Although freshwaters cover only a small fraction of the global surface, their contribution to global methane emission is significant and this is expected to increase, as a positive feedback to climate warming and exacerbated eutrophication. Yet, global estimates of methane emission from freshwaters are often based on point measurements that are spatio-temporally biased. To better constrain the uncertainties in quantifying methane fluxes from inland waters, a closer examination of the processes transporting methane from sediment to atmosphere is necessary. Among these processes, ebullition (bubbling) is an important transport pathway and is a primary source of uncertainty in quantifying methane emissions from freshwaters. This thesis aims to improve our understanding of ebullition in freshwaters by studying the processes of methane bubble formation, storage and release in aquatic sediments. The laboratory experiments demonstrate that aquatic sediments can store up to ~20% (volumetric content) gas and the storage capacity varies with sediment properties. The methane produced is stored as gas bubbles in sediment with minimal ebullition until the storage capacity is reached. Once the sediment void spaces are created by gas bubble formation, they are stable and available for future bubble storage and transport. Controlled water level drawdown experiments showed that the amounts of gas released from the sediment scaled with the total volume of sediment gas storage and correlated linearly to the drop in hydrostatic pressure. It was hypothesized that not only the timing of ebullition is controlled by sediment gas storage, but also the spatial distribution of ebullition. A newly developed freeze corer, capable of characterizing sediment gas content under in situ environments, enabled the possibility to test the hypothesis in a large subtropical lake (Lake Kinneret, Israel). The results showed that gas content was variable both vertically and horizontally in the lake sediment. Sediment methane production rate and sediment characteristics could explain these variabilities. The spatial distribution of ebullition generally was in a good agreement with the horizontal distribution of depth-averaged (surface 1 m) sediment gas content. While discrepancies were found between sediment depth-integrated methane production and the snapshot ebullition rate, they were consistent in a long term (multiyear average). These findings provide a solid basis for the future development of a process-based ebullition model. By coupling a sediment transport model with a sediment diagenetic model, general patterns of ebullition hotspots can be predicted at a system level and the uncertainties in ebullition flux measurements can be better constrained both on long-term (months to years) and short-term (minutes to hours) scales.
Periphyton-grazer interactions in headwater streams (2019)
Mewes, Daniela
Small headwater streams comprise most of the total channel length and catchment area in fluvial networks. They are tightly connected to their catchments and, thus, are highly vulnerable to changes in catchment hydrologic budgets and land use. Although these small, often fishless streams are of little economic interest, they are vital for the ecological and chemical state of larger water bodies. Although numerous studies investigate the impact of various anthropogenic stressors or altered catchment conditions, we lack an in-depth understanding of the natural conditions and processes in headwater streams. This natural state, however, largely affects how a headwater stream responds to anthropogenic or climatic changes. One of the major threats to aquatic ecosystems is the excessive anthropogenic input of nutrients leading to eutrophication. Nutrients exert a bottom-up effect in the food web, foremost affecting primary producers and their consumers, i.e. periphyton and benthic grazers in headwater streams. The periphyton-grazer link is the main path of autochthonous (in-stream) production into the stream food web and the strength of this link largely determines the effectiveness of this pathway. Therefore, this thesis aims at elucidating important biological processes with the explicit focus on periphyton-grazer interactions. I assessed different aspects of periphyton-grazer interactions using laboratory experiments to solve methodological problems, and using a field study to compare the benthic communities of three morphologically similar, phosphorus-limited, near-natural headwater streams. With the results of the laboratory experiments, I was able to show that periphyton RNA/DNA ratios can be used as proxy for periphyton growth rates in controlled experiments and that the fatty acid composition of grazing mayfly nymphs responds to changes in fatty acids provided by the diet after only two weeks. The use of the RNA/DNA ratio as a proxy for periphyton growth rate allows a comparison of these growth rates even in simple experimental set-ups and thereby permits the inclusion of this important process in ecotoxicological or ecological experiments. The observed fast turnover rates of fatty acids in consumer tissues show that even short-term changes in available primary producers can alter the fatty acid composition of primary consumers with important implications for the supply of higher trophic levels with physiologically important polyunsaturated fatty acids. With the results of the field study, I revealed gaps in the understanding of the linkages between catchment and in-stream phosphorus availability under near-natural conditions and demonstrated that seemingly comparable headwater streams had significantly different benthic communities. These differences most likely affect stream responses to environmental changes.
„… weil jeder Mensch gleich ist – im Prinzip.“ (2019)
Moser, Irene
Within the framework of a participatory action research approach, teachers and their students were accompanied scientifically with a focus on Montessori pedagogy and music in an upper secondary grammar school for three years (from 2012-2015). Based on attitude- and self-efficacy-theories, inclusive instruction as well as agent-based research, the aim was to examine the implementation of an inclusive learning - and school-culture at this grammar school – an institution which is part of the highly selective education system at secondary II level in Austria. It has been possible to reconstruct the understanding of teachers concerning inclusion and conceptions of man as well as their selfefficacy of competences to teach in an inclusive environment by means of qualitative and quantitative data. To examine attitudes and selfefficacy of teacher competences, international and national tested scales SACIE-R (Sentiments, Attitudes and Concerns about Inclusive Education-Revised) and TEIP (Teacher Efficacy for Inclusive Practice) were used in a German version. Furthermore, the empirical survey was extended by a webbased national questionnaire for teachers in different types of schools, from primary to secondary level (n= 427) to allow comparative analysis. Interviews, which were evaluated content-analytically and ques-tionnaires for parents (n=37) and students (n=19) offered an indepth insight into everyday educational practice from different perspectives. The synopsis of the result shows, that it was possible for a rather young team of teachers with very positive attitudes towards inclusion, to implement a supportive and vivid learning - and school culture. Nevertheless, the efforts of the teachers did not exceed the system limits. The development towards a ‘school for all’, a logical consequence of theoretical approaches concerning inclusion, was not discovered. The current research is verifying the corefindings of inclusive - and agent-based research. Further findings can be related to the selective structures of the education system in secondary II.
Aspects of Information Access: Modeling Navigation on Wikipedia (2018)
Dimitrov, Dimitar
Navigation is a natural way to explore and discover content in a digital environment. Hence, providers of online information systems such as Wikipedia---a free online encyclopedia---are interested in providing navigational support to their users. To this end, an essential task approached in this thesis is the analysis and modeling of navigational user behavior in information networks with the goal of paving the way for the improvement and maintenance of web-based systems. Using large-scale log data from Wikipedia, this thesis first studies information access by contrasting search and navigation as the two main information access paradigms on the Web. Second, this thesis validates and builds upon existing navigational hypotheses to introduce an adaptation of the well-known PageRank algorithm. This adaptation is an improvement of the standard PageRank random surfer navigation model that results in a more "reasonable surfer" by accounting for the visual position of links, the information network regions they lead to, and the textual similarity between the link source and target articles. Finally, using agent-based simulations, this thesis compares user models that have a different knowledge of the network topology in order to investigate the amount and type of network topological information needed for efficient navigation. An evaluation of agents' success on four different networks reveals that in order to navigate efficiently, users require only a small amount of high-quality knowledge of the network topology. Aside from the direct benefits to content ranking provided by the "reasonable surfer" version of PageRank, the empirical insights presented in this thesis may also have an impact on system design decisions and Wikipedia editor guidelines, i.e., for link placement and webpage layout.
Zum Kompetenzzuwachs in und durch Sport- und Bewegungsangebote an rheinland-pfälzischen Ganztagsschulen (2018)
Krampe, Christian
Im Fokus der Arbeit steht die Frage des Kompetenzzuwachses in und durch außerunterrichtliche Sport- und Bewegungsangebote an rheinland pfälzischen Ganztagsschulen, die von Trainerinnen und Trainern angeboten und durchgeführt werden. Um dieser Fragestellung nachzugehen, wurden zunächst Schul- und Unterrichtsqualitätsmodelle auf das außerunterrichtlichen Sport- und Bewegungsangebot übertragen, um das zu untersuchende Setting im Kontext der rheinland-pfälzischen Ganztagsschullandschaft zu verorten und mögliche Einflussfaktoren und Wechselwirkungen vor dem Hintergrund des Kompetenzzuwachses der Teilnehmerinnen und -nehmer aufzuzeigen. Dem nachgestellt wurden während der Darlegung theoretischer Vorüberlegungen, wie z.B. Aspekte zum außerschulischen Bildungsverständnis und Sport und Bewegung in den Erziehungswissenschaften, Kategorien und Kriterien ermittelt, deren Nachweis den Kompetenzzuwachs der Schülerinnen und Schüler stark vermuten lassen. Die Untersuchungsleitlinie dieser Arbeit wurde ein qualitatives Forschungsdesign eingebunden. Die herangezogene Forschungsstrategie bildet eine vergleichende Fallanalyse. Als Erhebungsmethode dient das Experteninterview; ein problemorientiertes Interview im Übungsleiterinnen und -leitern, die für die Planung und Durchführung der in rheinland-pfälzischen Ganztagsschulen implementierten Sport- und Bewegungsangebote verantwortlich zeichnen. Die Auswertung erfolgte über eine modifizierte Form der qualitativen Inhaltsanalyse. In der Summe konnte aufgezeigt werden, dass es sich bei den untersuchten Sport- und Bewegungsangeboten um hochwertige Settings handelt, in denen Kriterien und hypothetische Annahmen aus den Schul- und Unterrichtsqualitätsmodellen aufgezeigt werden konnten und somit günstige Voraussetzungen –die aktive Teilnahme vorausgesetzt– für den Kompetenzerwerb der Schülerinnen und Schüler an Ganztagsschulen vorliegen.
Bewertung geometrischer und radiometrischer Effekte digitaler Flächensensoren in der Bündeltriangulation (2018)
Kisser, Waldemar
Technologische Fortschritte auf dem Gebiet der integrierten Halbleitertechnik, die unter anderem auch zur gestiegenen Leistungsfähigkeit der Kamerasensoren beitragen, konzentrierten sich bisher primär auf die Schnelligkeit und das Auflösungsvermögen der Sensoren. Die sich ständig verändernde Entwicklung hat jedoch direkte Folgen auf das physikalische Verhalten einer Kamera und damit auch Konsequenzen für die erreichbare geometrische Genauigkeit einer photogrammetrischen 3D-Rekonstruktion. Letztere stand bisher nicht im Fokus der Forschung und ist eine Aufgabe, der sich diese Arbeit im Sinne der Photogrammetrie und Messtechnik stellt. Aktuelle Untersuchungen und Erfahrungen aus industriellen Projekten zeigen in diesem Zusammenhang, dass das geometrisch-physikalische Verhalten digitaler Kameras - für höchste photogrammetrische Ansprüche - noch nicht ausreichend modelliert ist. Direkte Aussagen zur erreichbaren Genauigkeit bei gegebener Hardware erweisen sich daher bislang als unzureichend. Ferner kommt es aufgrund der unpräzisen Modellierung zu Einbußen in der Zuverlässigkeit der erreichten Ergebnisse. Für den Entwickler präziser kamerabasierter Messverfahren folgt daraus, dass zu einer optimalen Schätzung der geometrischen Genauigkeit und damit auch vollständigen Ausschöpfung der Messkamera geeignete mathematische Modelle erforderlich sind, die das geometrisch physikalische Verhalten bestmöglich beschreiben. Diese Arbeit beschreibt, wie die erreichbare Genauigkeit einer Bündelblockausgleichung, schon a priori mithilfe des EMVA1288 Standards approximiert werden kann. Eine in diesem Zusammenhang wichtige Teilaufgabe ist die Schaffung einer optimalen Messanordnung. Hierzu gehören Untersuchungen der üblicherweise verwendeten Kalibrierkörper und die Beseitigung von systematischen Fehlern vor und nach der Bündelblockausgleichung. Zum Nachweis dieser Systematiken wird eine auf statistischem Lernen basierende Methode beschrieben und untersucht. Erst wenn alle genauigkeitsmindernden Einflüsse berücksichtigt sind, wird der Anteil des Sensors in den Messdaten sichtbar und damit auch mathematisch parametrisierbar. Die Beschreibung des Sensoreinflusses auf die erreichbare Genauigkeit der Bündelblockausgleichung erfolgt in drei Schritten. Der erste Schritt beschreibt den Zusammenhang zwischen ausgewählten EMVA1288-Kennzahlen und der Unsicherheit eines Grauwertes. Der zweite Schritt ist eine Modellierung dieser Grauwertunsicherheit als Zentrumsunsicherheit einer Zielmarke. Zur Beschreibung dieser Unsicherheit innerhalb der Bündelblockausgleichung wird ein stochastisches Modell, basierend auf dem EMVA1288-Standard, vorgeschlagen. Ausgehend vom Rauschen des Zielmarkenmittelpunktes wird im dritten Schritt die Unsicherheit im Objektraum beispielhaft mit Hilfe von physikalisch orientierten Simulationen approximiert. Die Wirkung der vorgeschlagenen Methoden wird anhand von Realkalibrierungen nachgewiesen. Abschließend erfolgt die Diskussion der vorgeschlagenen Methoden und erreichten Ergebnisse sowie ein Ausblick auf kommende Untersuchungen.
Automatische Klassifikation von hämatopoetischen Zellen für ein computer-assistiertes Mikroskopiesystem (2018)
Krappe, Sebastian
The cytological examination of bone marrow serves as clarification of variations in blood smears. It is also used for the clarification of anemia, as exclusion of bone marrow affection at lymphoma and at suspicion of leukemia. The morphological evaluation of hematopoietic cells is the basis for the creation of the diagnosis and for decision support for further diagnostics. Even for experienced hematologists the manual classification of hematopoietic cells is time-consuming, error-prone and subjective. For this reason new methods in the field of image processing and pattern recognition for the automatic classification including preprocessing steps are developed for a computer-assisted microscopy system. These methods are evaluated by means of a huge reference database. The proposed image analysis procedures comprise methods for the automated detection of smears, for the determination of relevant regions, for the localization and segmentation of single hematopoietic cells as well as for the feature extraction and classification task. These methods provide the basis for the first system for the automated, morphological analysis of bone marrow aspirates for leukemia diagnosis and are therefore a major contribution for a better and more efficient patient care in the future.
Retrospektive Analyse der Ausbreitung und dynamische Erkennung von Web-Tracking durch Sandboxing (2018)
Wambach, Tim
Aktuelle quantitative Analysen von Web-Tracking bieten keinen umfassenden Überblick über dessen Entstehung, Ausbreitung und Entwicklung. Diese Arbeit ermöglicht durch Auswertung archivierter Webseiten eine rückblickende Erfassung der Entstehungsgeschichte des Web-Trackings zwischen den Jahren 2000 und 2015. Zu diesem Zweck wurde ein geeignetes Werkzeug entworfen, implementiert, evaluiert und zur Analyse von 10000 Webseiten eingesetzt. Während im Jahr 2005 durchschnittlich 1,17 Ressourcen von Drittparteien eingebettet wurden, zeigt sich ein Anstieg auf 6,61 in den darauffolgenden 10 Jahren. Netzwerkdiagramme visualisieren den Trend zu einer monopolisierten Netzstruktur, in der bereits ein einzelnes Unternehmen 80 % der Internetnutzung überwachen kann. Trotz vielfältiger Versuche, dieser Entwicklung durch technische Maßnahmen entgegenzuwirken, erweisen sich nur wenige Selbst- und Systemschutzmaßnahmen als wirkungsvoll. Diese gehen häufig mit einem Verlust der Funktionsfähigkeit einer Webseite oder mit einer Einschränkung der Nutzbarkeit des Browsers einher. Mit der vorgestellten Studie wird belegt, dass rechtliche Vorschriften ebenfalls keinen hinreichenden Schutz bieten. An Webauftritten von Bildungseinrichtungen werden Mängel bei Erfüllung der datenschutzrechtlichen Pflichten festgestellt. Diese zeigen sich durch fehlende, fehlerhafte oder unvollständige Datenschutzerklärungen, deren Bereitstellung zu den Informationspflichten eines Diensteanbieters gehören. Die alleinige Berücksichtigung klassischer Tracker ist nicht ausreichend, wie mit einer weiteren Studie nachgewiesen wird. Durch die offene Bereitstellung funktionaler Webseitenbestandteile kann ein Tracking-Unternehmen die Abdeckung von 38 % auf 61 % erhöhen. Diese Situation wird durch Messungen von Webseiten aus dem Gesundheitswesen belegt und aus technischer sowie rechtlicher Perspektive bewertet. Bestehende systemische Werkzeuge zum Erfassen von Web-Tracking verwenden für ihre Messung die Schnittstellen der Browser. In der vorliegenden Arbeit wird mit DisTrack ein Framework zur Web-Tracking-Analyse vorgestellt, welches eine Sandbox-basierte Messmethodik verfolgt. Dies ist eine Vorgehensweise, die in der dynamischen Schadsoftwareanalyse erfolgreich eingesetzt wird und sich auf das Erkennen von Seiteneffekten auf das umliegende System spezialisiert. Durch diese Verhaltensanalyse, die unabhängig von den Schnittstellen des Browsers operiert, wird eine ganzheitliche Untersuchung des Browsers ermöglicht. Auf diese Weise können systemische Schwachstellen im Browser aufgezeigt werden, die für speicherbasierte Web-Tracking-Verfahren nutzbar sind.
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